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IONIS PHARMACEUTICALS INC
IONIS PHARMACEUTICALS INC. Institutional holdings reported to the SEC.

Institutional Holders (2017Q2)
Portfolio Summary: 315 Institutional holders with a total value of $5,772,701,525.
#
Manager Name
Shares
Value ($)
%
Options
Notes
51 SPOT TRADING LLC /BD 738 38,000 0.00%
52 BB&T INVESTMENT SERVICES, INC. 800 41,000 0.00%
53 First Horizon Advisors, Inc. 926 44,000 0.00%
54 Cornerstone Advisors, Inc. 910 46,000 0.00%
55 HALL LAURIE J TRUSTEE 990 50,361 0.00%
56 Riggs Asset Managment Co. Inc. 1,000 51,000 0.00%
57 Portfolio Solutions, LLC 1,000 51,000 0.00%
58 Perkins Coie Trust Co 1,000 51,000 0.00%
59 Winfield Associates, Inc. 1,010 52,000 0.00%
60 PARADIGM ASSET MANAGEMENT CO LLC 1,100 55,957 0.00%
61 HUNTINGTON NATIONAL BANK 1,100 56,000 0.00%
62 ETF MANAGERS GROUP, LLC 1,204 61,000 0.00%
63 BESSEMER GROUP INC 1,242 63,000 0.00%
64 Global X Management CO LLC 1,335 68,000 0.00%
65 CUTLER GROUP LLC / CA 28,800 77,000 0.00% Put
66 BALENTINE LLC 1,716 87,000 0.00%
67 FLORENCE CAPITAL ADVISORS, LLC 2,000 92,000 0.00%
68 JPMORGAN CHASE & CO 1,809 92,000 0.00%
69 KBC Group NV 1,869 95,000 0.00%
70 Kistler-Tiffany Companies, LLC 1,900 97,000 0.00%
71 STEWARD PARTNERS INVESTMENT ADVISORY, LLC 1,975 100,000 0.00%
72 YORKBRIDGE WEALTH PARTNERS, LLC 2,031 106,000 0.00%
73 PARALLEL ADVISORS, LLC 2,073 109,000 0.00%
74 SIMPLEX TRADING, LLC 2,631 133,000 0.00%
75 MOSAIC FAMILY WEALTH, LLC 2,635 134,000 0.00%
76 AMERICAN INTERNATIONAL GROUP, INC. 2,994 152,000 0.00%
77 HANSEATIC MANAGEMENT SERVICES INC 3,481 177,000 0.00%
78 MANUFACTURERS LIFE INSURANCE COMPANY, THE 3,744 190,000 0.00%
79 WFG Advisors, LP 3,800 193,000 0.00%
80 QS Investors, LLC 4,010 204,000 0.00%
81 SPOT TRADING LLC /BD 4,100 209,000 0.00% Call
82 Squarepoint Ops LLC 4,328 220,000 0.00%
83 REGIONS FINANCIAL CORP 4,321 220,000 0.00%
84 FORESTERS INVESTMENT MANAGEMENT COMPANY, INC. 4,361 222,000 0.00%
85 Cambridge Investment Research Advisors, Inc. 4,362 222,000 0.00%
86 JANE STREET GROUP, LLC 4,465 227,000 0.00%
87 PRELUDE CAPITAL MANAGEMENT, LLC 4,570 232,000 0.00%
88 Trexquant Investment LP 4,580 233,000 0.00%
89 Zurcher Kantonalbank (Zurich Cantonalbank) 4,880 248,000 0.00%
90 Cheyne Capital Management (UK) LLP 4,891 249,000 0.00%
91 EQUITEC SPECIALISTS, LLC 5,000 254,000 0.00% Put
92 CAPSTONE ASSET MANAGEMENT CO 5,018 255,000 0.00%
93 SOMERVILLE KURT F 5,079 258,000 0.00%
94 BANK OF MONTREAL /CAN/ 5,071 258,000 0.00%
95 US BANCORP \DE\ 5,152 262,000 0.00%
96 Private Advisor Group, LLC 5,102 263,000 0.00%
97 DUPONT CAPITAL MANAGEMENT CORP 5,184 264,000 0.00%
98 FDx Advisors, Inc. 5,272 268,000 0.00%
99 First City Capital Management, Inc. 5,330 271,000 0.00%
100 BLAIR WILLIAM & CO/IL 5,381 274,000 0.00%
Page 2 of 7

From 2023-01-03 onward, the SEC requires values to be reported in nearest dollars. Earlier filings reported values in thousands; therefore, positions shown as $0 in older filings may represent actual values between $0–$999.