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IONIS PHARMACEUTICALS INC
IONIS PHARMACEUTICALS INC. Institutional holdings reported to the SEC.

Institutional Holders (2017Q2)
Portfolio Summary: 315 Institutional holders with a total value of $5,772,701,525.
#
Manager Name
Shares
Value ($)
%
Options
Notes
101 WOLVERINE TRADING LP /BD 50,300 2,606,000 0.05% Call
102 LOS ANGELES CAPITAL MANAGEMENT LLC 51,135 2,601,000 0.05%
103 PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO 51,102 2,600,000 0.05%
104 D.A. DAVIDSON & CO. 50,957 2,592,000 0.04%
105 CITADEL ADVISORS LLC 50,300 2,559,000 0.04% Put
106 NJ State Employees Deferred Compensation Plan 50,000 2,543,000 0.04%
107 VOYA INVESTMENT MANAGEMENT LLC 47,737 2,428,000 0.04%
108 PROFUND ADVISORS LLC 47,274 2,405,000 0.04%
109 ENVESTNET ASSET MANAGEMENT INC 45,500 2,314,000 0.04%
110 Legal & General Group Plc 44,480 2,262,000 0.04%
111 Opus Point Partners Management, LLC 43,867 2,232,000 0.04%
112 WOLVERINE TRADING LP /BD 41,900 2,170,000 0.04% Put
113 First Light Asset Management, LLC 42,015 2,137,000 0.04%
114 CITIGROUP INC 41,700 2,121,000 0.04% Call
115 CAPITAL COUNSEL LLC/NY 41,280 2,100,000 0.04%
116 ROYAL BANK OF CANADA 40,700 2,070,000 0.04% Put
117 GOLDMAN SACHS GROUP INC 37,700 1,918,000 0.03% Put
118 GOLDMAN SACHS GROUP INC 37,700 1,918,000 0.03% Call
119 TRAIL RIDGE CAPITAL LLC 31,366 1,596,000 0.03%
120 BANK OF NOVA SCOTIA 30,000 1,526,000 0.03%
121 Neuberger Berman Group LLC 29,412 1,496,000 0.03%
122 TWO SIGMA ADVISERS, LP 29,111 1,481,000 0.03%
123 Paloma Partners Management Co 28,500 1,450,000 0.03%
124 Employees Retirement System of Texas 28,000 1,424,000 0.02%
125 CHICAGO EQUITY PARTNERS LLC 27,770 1,413,000 0.02%
126 BlueCrest Capital Management Ltd 27,706 1,409,000 0.02%
127 Creative Planning 26,499 1,348,000 0.02%
128 Virtu KCG Holdings LLC 26,386 1,342,000 0.02%
129 SIT INVESTMENT ASSOCIATES INC 25,675 1,306,000 0.02%
130 BTG Pactual Global Asset Management Ltd 25,388 1,291,000 0.02%
131 WOLVERINE ASSET MANAGEMENT LLC 25,000 1,271,000 0.02% Call
132 STATE OF WISCONSIN INVESTMENT BOARD 24,616 1,252,000 0.02%
133 PUBLIC EMPLOYEES RETIREMENT ASSOCIATION OF COLORADO 24,390 1,241,000 0.02%
134 AMALGAMATED BANK 23,191 1,180,000 0.02%
135 J.P. Morgan Private Wealth Advisors LLC 23,017 1,171,000 0.02%
136 Utah Retirement Systems 22,600 1,149,000 0.02%
137 RELIANCE TRUST CO OF DELAWARE 22,086 1,124,000 0.02%
138 GSA CAPITAL PARTNERS LLP 21,545 1,096,000 0.02%
139 OPPENHEIMER ASSET MANAGEMENT INC. 21,502 1,094,000 0.02%
140 Raymond James Financial Services Advisors, Inc. 21,475 1,092,000 0.02%
141 PRINCIPAL FINANCIAL GROUP INC 21,445 1,091,000 0.02%
142 BB&T SECURITIES, LLC 21,055 1,071,000 0.02%
143 VIRTUS ETF ADVISERS LLC 20,375 1,036,000 0.02%
144 RUSSELL INVESTMENTS GROUP, LTD. 20,077 1,021,000 0.02%
145 FIC CAPITAL INC 19,290 981,000 0.02%
146 COMERICA BANK 16,171 963,000 0.02%
147 Commonwealth Equity Services, Inc 18,876 960,000 0.02%
148 TD ASSET MANAGEMENT INC 18,800 956,000 0.02%
149 AMERIPRISE FINANCIAL INC 18,752 953,000 0.02%
150 WRAPMANAGER INC 18,577 945,000 0.02%
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From 2023-01-03 onward, the SEC requires values to be reported in nearest dollars. Earlier filings reported values in thousands; therefore, positions shown as $0 in older filings may represent actual values between $0–$999.