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LIGAND PHARMACEUTICALS INC
LIGAND PHARMACEUTICALS INC. Institutional holdings reported to the SEC.

Institutional Holders (2015Q4)
Portfolio Summary: 235 Institutional holders with a total value of $2,497,264,026.
#
Manager Name
Shares
Value ($)
%
Options
Notes
1 BlackRock Fund Advisors 1,353,035 146,696,000 5.87%
2 VANGUARD GROUP INC 1,348,453 146,199,000 5.85%
3 KNOTT DAVID M 1,199,766 130,078,630 5.21%
4 WILLIAM BLAIR INVESTMENT MANAGEMENT, LLC 1,140,511 123,654,000 4.95%
5 FMR LLC 1,110,563 120,407,000 4.82%
6 RS INVESTMENT MANAGEMENT CO LLC 1,089,730 118,149,000 4.73%
7 OAK RIDGE INVESTMENTS LLC 955,599 103,606,000 4.15%
8 CARDINAL CAPITAL MANAGEMENT LLC /CT 852,053 92,380,000 3.70%
9 Invesco Ltd. 740,076 80,239,000 3.21%
10 STATE STREET CORP 635,378 68,885,000 2.76%
11 DIMENSIONAL FUND ADVISORS LP 551,168 59,758,000 2.39%
12 WELLS FARGO & COMPANY/MN 549,426 59,568,000 2.39%
13 Bank of New York Mellon Corp 544,054 58,986,000 2.36%
14 PRICE T ROWE ASSOCIATES INC /MD/ 478,988 51,932,000 2.08%
15 BlackRock Institutional Trust Company, N.A. 473,441 51,330,000 2.06%
16 NORTHERN TRUST CORP 441,421 47,858,000 1.92%
17 RUSSELL FRANK CO/ 418,475 45,266,000 1.81%
18 DEUTSCHE BANK AG\ 391,562 42,451,000 1.70%
19 Stephens Investment Management Group LLC 389,298 42,208,000 1.69%
20 Granahan Investment Management, LLC 358,624 38,882,000 1.56%
21 AMERIPRISE FINANCIAL INC 323,985 35,126,000 1.41%
22 BANK OF AMERICA CORP /DE/ 302,858 32,836,000 1.31%
23 KALMAR INVESTMENTS INC /DE/ 293,141 31,782,000 1.27%
24 Conestoga Capital Advisors, LLC 286,162 31,026,000 1.24%
25 Allianz Asset Management GmbH 252,731 27,401,000 1.10%
26 MACQUARIE GROUP LTD 232,356 25,192,000 1.01%
27 NEXT CENTURY GROWTH INVESTORS LLC 226,892 24,599,000 0.99%
28 JANUS HENDERSON INVESTORS US LLC 207,214 22,466,000 0.90%
29 CASTLEARK MANAGEMENT LLC 206,576 22,397,000 0.90%
30 PANAGORA ASSET MANAGEMENT INC 198,597 21,532,000 0.86%
31 AJO, LP 188,594 20,447,000 0.82%
32 RICE HALL JAMES & ASSOCIATES, LLC 180,412 19,560,000 0.78%
33 RENAISSANCE TECHNOLOGIES LLC 174,300 18,898,000 0.76%
34 FIDUCIARY MANAGEMENT ASSOCIATES LLC 161,051 17,462,000 0.70%
35 EAGLE ASSET MANAGEMENT INC 159,301 17,271,000 0.69%
36 NORGES BANK 154,165 16,715,000 0.67%
37 GEODE CAPITAL MANAGEMENT, LLC 128,971 13,983,000 0.56%
38 RK Capital Management, LLC 127,983 13,876,000 0.56%
39 PRINCIPAL FINANCIAL GROUP INC 122,475 13,279,000 0.53%
40 CHARLES SCHWAB INVESTMENT MANAGEMENT INC 108,401 11,753,000 0.47%
41 ASHFORD CAPITAL MANAGEMENT INC 103,420 11,213,000 0.45%
42 UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC 102,784 11,143,000 0.45%
43 OBERWEIS ASSET MANAGEMENT INC/ 101,127 10,964,000 0.44%
44 GUGGENHEIM CAPITAL LLC 94,060 10,198,000 0.41%
45 Nichols Asset Management LLC 93,170 10,101,000 0.40%
46 MORGAN STANLEY 85,245 9,243,000 0.37%
47 PRUDENTIAL FINANCIAL INC 84,432 9,154,000 0.37%
48 CREDIT SUISSE AG/ 84,315 9,141,000 0.37%
49 AMERICAN CENTURY COMPANIES INC 82,441 8,938,000 0.36%
50 BELLEVUE ASSET MANAGEMENT AG 81,640 8,851,000 0.35%
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From 2023-01-03 onward, the SEC requires values to be reported in nearest dollars. Earlier filings reported values in thousands; therefore, positions shown as $0 in older filings may represent actual values between $0–$999.