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LIGAND PHARMACEUTICALS INC
LIGAND PHARMACEUTICALS INC. Institutional holdings reported to the SEC.

Institutional Holders (2016Q1)
Portfolio Summary: 256 Institutional holders with a total value of $2,710,947,783.
#
Manager Name
Shares
Value ($)
%
Options
Notes
1 VANGUARD GROUP INC 1,404,007 150,355,000 5.55%
2 BlackRock Fund Advisors 1,377,004 147,463,000 5.44%
3 WILLIAM BLAIR INVESTMENT MANAGEMENT, LLC 1,229,452 131,662,000 4.86%
4 KNOTT DAVID M 1,199,766 128,482,941 4.74%
5 FMR LLC 975,263 104,441,000 3.85%
6 OAK RIDGE INVESTMENTS LLC 941,493 100,825,000 3.72%
7 RS INVESTMENT MANAGEMENT CO LLC 912,545 97,724,000 3.60%
8 CARDINAL CAPITAL MANAGEMENT LLC /CT 839,691 89,923,000 3.32%
9 WELLS FARGO & COMPANY/MN 839,205 89,870,000 3.32%
10 1832 Asset Management L.P. 469,700 80,340,000 2.96%
11 STATE STREET CORP 699,788 74,940,000 2.76%
12 AJO, LP 599,815 64,234,000 2.37%
13 Bank of New York Mellon Corp 589,823 63,163,000 2.33%
14 Invesco Ltd. 571,509 61,204,000 2.26%
15 PRICE T ROWE ASSOCIATES INC /MD/ 490,928 52,573,000 1.94%
16 BlackRock Institutional Trust Company, N.A. 473,544 50,712,000 1.87%
17 RUSSELL FRANK CO/ 472,386 50,588,000 1.87%
18 DEUTSCHE BANK AG\ 451,355 48,334,000 1.78%
19 NORTHERN TRUST CORP 423,188 45,319,000 1.67%
20 Conestoga Capital Advisors, LLC 413,362 44,267,000 1.63%
21 AMERIPRISE FINANCIAL INC 389,327 41,693,000 1.54%
22 Stephens Investment Management Group LLC 386,266 41,365,000 1.53%
23 Granahan Investment Management, LLC 382,624 40,975,000 1.51%
24 DIMENSIONAL FUND ADVISORS LP 366,468 39,244,000 1.45%
25 NEXT CENTURY GROWTH INVESTORS LLC 314,860 33,718,000 1.24%
26 BANK OF AMERICA CORP /DE/ 308,824 33,073,000 1.22%
27 KALMAR INVESTMENTS INC /DE/ 282,071 30,207,000 1.11%
28 MACQUARIE GROUP LTD 270,913 29,012,000 1.07%
29 Allianz Asset Management GmbH 260,016 27,845,000 1.03%
30 CASTLEARK MANAGEMENT LLC 213,551 22,869,000 0.84%
31 JANUS HENDERSON INVESTORS US LLC 204,796 21,932,000 0.81%
32 PANAGORA ASSET MANAGEMENT INC 197,271 21,126,000 0.78%
33 PARAMETRIC PORTFOLIO ASSOCIATES LLC 194,183 20,795,000 0.77%
34 RICE HALL JAMES & ASSOCIATES, LLC 184,371 19,744,000 0.73%
35 FIDUCIARY MANAGEMENT ASSOCIATES LLC 160,716 17,208,000 0.63%
36 EAGLE ASSET MANAGEMENT INC 160,677 17,206,000 0.63%
37 COLUMBIA WANGER ASSET MANAGEMENT LLC 158,034 16,924,000 0.62%
38 NORGES BANK 154,165 16,510,000 0.61%
39 RK Capital Management, LLC 151,783 16,254,000 0.60%
40 GEODE CAPITAL MANAGEMENT, LLC 131,624 14,095,000 0.52%
41 MORGAN STANLEY 129,242 13,840,000 0.51%
42 PRINCIPAL FINANCIAL GROUP INC 127,403 13,643,000 0.50%
43 Summit Creek Advisors LLC 126,197 13,514,000 0.50%
44 RENAISSANCE TECHNOLOGIES LLC 116,800 12,508,000 0.46%
45 TIAA CREF INVESTMENT MANAGEMENT LLC 108,870 11,659,000 0.43%
46 ASHFORD CAPITAL MANAGEMENT INC 103,545 11,089,000 0.41%
47 FALCON POINT CAPITAL, LLC 102,263 10,951,000 0.40%
48 TEACHERS ADVISORS, LLC 100,617 10,775,000 0.40%
49 UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC 95,730 10,251,000 0.38%
50 CREDIT SUISSE AG/ 95,212 10,196,000 0.38%
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From 2023-01-03 onward, the SEC requires values to be reported in nearest dollars. Earlier filings reported values in thousands; therefore, positions shown as $0 in older filings may represent actual values between $0–$999.